WHAT ORCHID IS THIS? HOW PHOTOS CAN HELP! – Part Two of Two Parts

Part One covered hints for photographing orchids so as to be able to identify the plant.  Part Two gives an example with Cyrtostylis robusta (Winter Gnat Orchid) and C. reniformis (Small Gnat Orchid).

Although the flowering times are different – C reniformis is spring flowering and C. robusta is winter flowering – there is a slight overlap in August when it is possible for both to be flowering at the same time and in the same place.

The flowers are very similar but major difference between the two species is the leaf.  Both leaves are roughly kidney shape but C. robusta is pale green with pale , almost undistinguished veins, silvery underneath whilst  C. reniformis is heavily veined, blue-green above and green below.

Cyrtostylis reniformis (4)
C. reniformis, easily identified by the leaf, even when in bud.

The other differences are more subtle.

  • C. reniformis has dark buds and the apex of the labellum tends to be rounded rather than pointed
  • C. robusta has pale reddish buds and larger flowers, labellum crenulated (slightly wavy) and a fine point at the apex.
Cyrtostylis reniformis
C. reniformis – This is not a good identification picture because though the leaf is present, it is too blurry for identifying the species and the angle of the flower obscures the labellum  details

Consequently, it is important that photographs of the flowers clearly show the labellum – pointed labellum apex for C. robusta compared with the rounded labellum tip of C. reniformis.

 

Cyrtostylis robusta
In this photograph it is possible to identify the plants as C. robusta because of the leaves even though there are no clear views of the labellum.

March 2016 Winning Photograph

1603 sm JS Arachnorchis sp

We frequently receive entries from Western Australia but this month our entries were from both the west and the east. Allen Jennings entered a Calanthe triplicata (Christmas Orchid) from New South Wales. Pauline Meyer’s was from the west, (Western) Flying Duck Orchid, Paracaleana nigrita. The other entries were South Australian, Jenny Pauley’s recently photographed Leporella fimbriata (Fringed Hare Orchid), Greg Sara’s Thelymitra rubra (Common Pink Sun Orchid) and Judy Sara’s Plumatochilos sp. (Bearded Orchid) and Arachnorchis sp. (Spider Orchid).

The winning photograph was Judy’s Spider Orchid. Obviously it was one of the Green Comb Spider Orchids – A. dilatata complex. Of this group there are about a dozen possibilities. Knowing the location, Mt Boothby, helped to narrow the options with the most likely candidate being Arachnorchis stricta but it wasn’t convincing. It would appear that the tips of the sepals may have been chewed off when in bud.

A distinguishing feature of this species is that there are no clubs or osmophores on the sepals. Other species of this complex have clubs. Another feature is that the dorsal sepal is bent over the column unlike many other green combs which have an erect dorsal sepal. The features that caused doubt were lateral sepals looking droopy instead of being characteristically stiffly held out but dry conditions could cause this. The other was that the labellum did not strictly fit the description of A. stricta but then again it is a variable species.

The conclusion was a possible hybrid but there is no information on the likely parents or that is an atypical A. stricta that may have been damaged in bud.

This is an example of the difficulties that can occur when attempting to identify a plant from one photograph.

Reference:

Personal communications Thelma Bridle (NOSSA Conservation Officer)

Personal communications Bob Bates

Bates, R. J., ed. (2011). South Australian Native Orchids. Electronic version, 2011. NOSSA

Rules of entry:

The subject matter must have something to do with Australian orchids.  Any format is acceptable including Photo shopped images, artwork, etc

How to enter:

Email nossa.enquiries@gmail.com – jpg as large as you are able to send it, preferably A4 ratio for printing

Post: PO Box 565, Unley, 5061

Bring in to the meeting

WHAT ORCHID IS THIS? HOW PHOTOS CAN HELP! – Part One of Two Parts

The following article is from the April 2016 Journal of the Native Orchid Society of South Australia.  The article is complete in itself but Part Two of this post will illustrate how images can help with images of Crytostylis robusta and C. reniformis.

Orchids are beautiful plants and many of us like to capture that beauty on photographs. And there are many beautiful pictures around.

Many times NOSSA, the Herbarium and other specialist groups receive images requesting identification but the vital information is missing.

When photographing for identification, it is necessary to take more than one image, particularly if you are unable to easily return to the site for more images. When in doubt, take several shots from many different angles highlighting different features of the plant and its habitat.

Another very important point to remember is, when there are several of the same plant, to photograph the orchid that is most representative of the group, not the atypical or unusual plant.

General Guide

As a general guide, it is helpful to take a picture of each of the following

  • whole plant
  • individual flower – both from the front and the side, occasionally the back.
  • flower head
  • leaf or leaves
  • habitat

Other helpful things to consider photographing are:

  • capsules of the finished flower – sometimes it can yield useful information.
  • for some genera, the stem can also be a helpful feature as between some species there can be a difference in the hairiness of the stem.
  • It is also worth including in a photograph an indication of whether the plants are growing in colonies with others or as scattered individual plants.

Importance of Size

It is also good to give an idea of size, this can be as simple as using a thumb or hand, a coin (show the reverse not the head) or any item that had an easily recognized size. It is important to have the object next to the feature being photographed. For example, a coin on the ground next to a leaf or a hand immediately behind the flower gives a clear indication of size. Remember to take another photo without the hand or coin.

Some Specific Identifying Features

Some species are distinctive and easily recognised, eg the Flying Duck Orchid, but others are not and it is helpful to know what part of the plant to photograph as different genera will have different identifying features.

  • Spider orchids – the tips of the segments (petals and sepals) and details of the labellum are important
  • Sun orchids – the column in the middle, the ovary at the base of the flower, and the number of bracts (leaf-like growth) on the stems
  • Pink fingers – the length of the leaf in comparison with the length of the flower stalk; also the back of the flower is helpful
  • Gnat orchid – the leaf is the best identifying feature, but also the bud can be helpful
  • Hyacinth orchid (of the Adelaide Hills) – labellum
  • Mosquito, Mayfly and Helmet orchids when not in flower – both sides of the leaf
  • Gastrodia – the flower spike
  • Rufoushoods – side view of the flower and close up of the labellum as the hairs on or surrounding the labellum are important features.
  • Leek Orchids – the labellum is very important, as well as part if not all of the flower spike as the distance between the individual flowers aids identification
  • Greenhoods – if present, the non-flowering rosette of leaves

The Australian Virtual Herbarium has some good tips for photographing flower. Click here to visit the site

 

Spider Orchid
Spider Orchid Photo: Robert Lawrence

This image lacks the ends of the segments to determine the identification.  The presence or absence of  clubs on the ends of the segments (petals and sepals) can often be the distinguishing feature with many of the Arachnorchis (Spider Orchids).

Atypical Cyrtostylis leaves

These leaves are an unusual shape and by themselves are not suitable for identification

CLUES TO ORCHID IDENTIFICATION – Prasophyllum & Microtis Leaves

Orchids are attractive and abound in variety. It is the variety that often provides the challenge of identification. As a novice it can be a bit overwhelming. In the eye of the beginner, the experienced orchid hunters appear to have no difficulty with identification. Over the years they have accumulated various clues that guide them toward accurate identification.

This series aims to document the clues that orchid hunters use.

Prasophyllum and Microtis Leaves

The first in the series relates to distinguishing between Microtis and Prasophyllum leaves. When in flower it is easy to see which is which but not so when only in leaf; and as they do not always produce flowers it is helpful to be able to separate them out at leaf stage.

Both leaves are green. Both are cylindrical. Both are hollow. Both resemble onion leaves.

M & P Leaves
Both types of leaves are long, thin and green.  The damaged Microtis leaf (below) shows the hollow.

The differences can be found in one or two areas. Microtis leaves are always green at the base whereas Prasophyllum leaves usually but not always will have a red or purplish coloured base. To help in identification, it is necessary to examine the base by moving the leaf litter aside to see where the plant emerges from the soil.

M & P Leaf Bases
The bases reveal the difference between the two genera.

Prasophyllum species that could have a green base are P. laxum, P. occulatans, P. sp Jip Jip, P. elatum, P. sp Sandplain, P. pallidum (although this is short and usually in bud when noticed), P. spicatum, P validum.  So further observations are necessary.

Another other area of difference is that the broken leaf of a Microtis yields a mucilaginous (thick sticky) sap; the Prasophyllum leaf does not.

Within the segregate genera used by many NOSSA members, there are two other genera with similar leaves. They are Microtidium and Hydrorchis. Again they are green, cylindrical, hollow but only hollow in the lower half. The top half is solid.

It should also be noted that Microtis can form dense colonies but Prasophyllum will never form more than loose colonies.

Finally, if upon gently feeling the base of these leaves it feels solid, that will indicate that there is a bud and it will most likely flower this season.

M & P Flowers
Once in flower, the differences between the two genera is obvious.

September 2015 Winning Photograph

09 sm PM A ferruginea possible
Caladenia huegelii complex
Plumatichlos sp Woodland Bearded Greenhood
Plumatichlos sp Woodland Bearded Greenhood

Five photos were entered for the September competition and there was a draw, Pauline Meyers’ flower of a plant from the Caladenia hueguelii complex from Western Australia and Jill McPherson’s Plumatichilos sp. Woodland Bearded Greenhood from Scott Creek Conservation Park. The other three by Chris Davey were photographed on Yorke Peninsula (see Letter to the Editor in this September Journal).
As winners they showcase the great diversity that are found in our Australian terrestrial orchids.

Plumatichilos belongs to the greenhoods but the features that set it apart from the other greenhoods are the long thin bristled labellum, the galea pinched in the middle resulting in two openings and the rosette of leaves growing a short way up the stem. Nationally there are thought to be several species but only a small handful have been named. In South Australia, there may be a few distinct species but currently they are usually identified with a phrase name such as Plumatichilos sp Woodland Bearded Greenhood.

Mainly flowering in spring the flowers of the Caladenia huegelii complex are characterised by the thickened clubs on the three sepals, petals shorter than the long sepals, fringed (either short or long) labellum with four or more rows of calli. All of these features can be seen in Pauline’s photograph but the leaf is not so easily seen which should be long, hairy and curve inward. In all there are said to be twenty two species within this complex of which twenty are named.

References:
Brown et al (2013) Field Guide to the Orchids of Western Australia, Floreat, WA Simon Nevill Publications.
Jones, David L (2006) A complete Guide to Native Orchids of Australia, Frenchs Forest, NSW: Reed New Holland
Jeans, Jeffrey & Gary Backhouse (2006) Wild Orchids of Victoria, Seaford Vic: Aquatic Photographics.
Bates, R. J. (2011) South Australia’s Native Orchids NOSSA DVD, Adelaide

Clues to Orchid Identification – Columns

Orchids can be incredibly easy to identify or frustratingly difficult.  For instance, the Flying Duck orchid is easily recognizable.  The name says it all. But for many other orchids, particularly the ubiquitous blue sun orchids, it is necessary to make careful observations.

 

 

Which is which
In this collage there are several species, but which is which?

 

For sun orchids the most important distinguishing feature will often be the column, details of which are described or illustrated in orchid keys.  For example Bates and Weber, Orchids of South Australia (1990) has an illustrated dichotomous keys showing the different types of columns.

As part of the key, drawings of the column were included
Images from Bates & Weber, Orchids of South Australia, pages 147 to 150; courtesy of the Board of the Botanic Gardens and State Herbarium.

With today’s cameras, it is possible to photograph the column, preferably from front, side and above.  This helps greatly with identification.

The following photographs illustrate the variety of columns.

Thelymitra alcockiae
Thelymitra alcockiae

 

Thelymitra grandiflora
Thelymitra brevifolia
Thelymitra brevifolia
Thelymitra cyanea
Thelymitra cyanea
Thelymitra peniculata
Thelymitra peniculata
Thelymitra pallidifructus
Thelymitra pallidifructus
Thelymitra juncifolia
Thelymitra juncifolia
Thelymitra inflata
Thelymitra inflata
Thelymitra arenaria
Thelymitra arenaria

 

Thelymitra megcalyptra
Thelymitra megcalyptra
Thelymitra X truncata - a natural hybrid
Thelymitra X truncata – a natural hybrid

And it is not only the blue orchids that can be a problem, in South Australia; there are three pink orchids which are different in size and flowering time, features that are not obvious in a photograph,  so the column is a helpful identification tool.

Thelymitra luteocilium
Thelymitra luteocilium – has tufted hairy column arms
Thelymitra rubra
Thelymitra rubra – has finger-like column arms and no tuft
Thelymitra carnea
Thelymitra carnea – has smooth column arms

 

And just what is the column? It is the reproductive organ of the orchid flower and is unique amongst plants because it is the fusion of the male (stamens) and female (pistil) parts.  It is usually found in the centre of the flower and both releases and receives the pollen sac.

 

 

 

 

 

 

2015 July Winning Photograph

07 sm JP Anzybas unguiculatus 2

Anzybas unguiculatas (common name Little Pelican or Cherry Helmet Orchid) was the main focus for this month’s competition with three photographs of this diminutive flower.

The other photographs were Ed Lowrey’s close-up of a triggered Urochilus sanguineus labellum, John Badger’s first Diuris palustris sighting for this year and Pauline Meyer’s mass flowering of Leptoceras menziesii post fire. Of the Anzybas, Jenny Pauley entered two and Lorraine Badger one. Jenny Pauley’s photograph of two flowers was the outstanding winner.

Originally named Corysanthes unguiculata (1810), then Corybas unguiculatus (1871), the genus name was changed in 2002 to Anzybas in recognition of its distribution both in Australia in New Zealand. Since 1945 it had been recognised that the New Zealand species Corybas cheesemanii was a synonym for Corybas unguiculatas although the juvenile plant can have two leaves unlike the Australian species which is single leafed.

An interesting feature of this flower is the prominent white ears at the rear of the helmet (not clearly seen in this photograph) which are part of the labellum.

These plants are small. The gum leaves and twigs give an idea of size but the engagement ring shows it very clearly.
These plants are small. The gum leaves and twigs give an idea of size but the engagement ring shows it very clearly.  Note also the prominent ‘white ears’ of the labellum.

An unusual aspect of this photograph is that the colour of the underside of one of the leaves. It lacks the characteristic distinguishing feature of the purple underside of the leaves. According to orchid growers, the light affects the leaf colour. Heavy shade produces green leaves. It is possible that the heavy leaf litter where this plant was growing provided enough deep shade to cause the colour loss.

Bates (1990) states that it (has) not proved amenable to cultivation, but it has, on rare occasions, been benched at NOSSA meetings with the most recent occurrence was in July 2010 but it remains a very difficult plant to cultivate. The electronic version Vol 34 No 7 has a photograph of the plant just visible within the moss.

It is not always easy to photograph this species as not only is it rare with limited numbers but there are very few sites where it can be found. Added to that is that the window of opportunity is short in South Australia with a flowering time from June to August compared with those interstate which can range from May to October.

There has always been an interest in Australian orchids.  Over the years there have been many photographs of orchids.  This stereographic postcard from 1928 is a study in beauty – https://i0.wp.com/www.slv.vic.gov.au/pcards/0/0/4/im/pc004332.jpg

This postcard is held by the State Library of Victoria.

Although the distribution covers the Southern Lofty, Kangaroo Island and the South East regions of South Australia, it has become increasingly rare due to loss of habitat which consists of leaf litter on damp soils.  As a result, there are very limited localities where they can now be found.

It is one of our earliest helmets to flower which is from June to July.

References

http://www.flora.sa.gov.au/cgi-bin/speciesfacts_display.cgi?form=speciesfacts&family=&genus=corybas&species=unguiculatus&iname=&submit=Display accessed August 5 2015

Les Nesbitt personal communication

Jones, David L (2006) Native Orchids of Australasia, Frenchs Forest, NSW: Reed New Holland.

Bates, R.J. & and Weber, J.Z. (1990). Orchids of South Australia, Adelaide: Flora and Fauna of S.A. Handbooks Committee

https://biodiversity.org.au/nsl/services/instance/apni/499184 accessed August 5 2015

https://www.anbg.gov.au/cpbr/cd-keys/orchidkey/html/genera/Anzybas.htm accessed August 5 2015

Bates R, (Editor) 2011 South Australia’s Native Orchids (CD-ROM), Adelaide: NOSSA

Journal of the Native Orchid Society of South Australia Vol 34 No 7 August 2010

http://rsnz.natlib.govt.nz/volume/rsnz_76/rsnz_76_04_007280.html accessed August 5 2015

Rupp, HMR and Hatch, ED (1945) Relation of the Orchid Flora of Australia to that of New Zealand in Proceeding of the Linnean Society of New South Wales Vol 70 1945, pages 53 – 61

https://archive.org/stream/proceedingsoflin70linn#page/60/mode/2up accessed August 5 2015

Orchid Identification Quiz #1

In 2006 the electronic version of the Journal of the Native Orchid Society of South Australia Volume 30 No 2 March contained a quiz – Do you Know Your Leaves? It featured photographs of the leaves of 15 different orchid. In 2011 Robert Lawrence produced a book titled Start With the Leaves which was based upon the premise that the flower is not always present but identification is still possible.

Many but not all can be identified to species level by the leaf alone.

So here is the web version of the original quiz – how well do you know your leaves?

Hint – 20 of them are South Australian, more specifically, the Mount Lofty Ranges region.  One is a weed.

Arachnorchis tentaculataThelymitra benthamiana Thelymitra antennifera Pyrorchis nigricans Plumatochilus sp Woodland Bearded Greenhood  Oligochaetochilus bisetusOrthoceras strictum Nemacianthus caudatus Eriochilus sp Hills Woodland  Leporella fimbriata Diuris palustrisGlossodia major

Diuris orientis

Diuris behrii

Cyrtostylis reniformisDisa bracteata - weed  Corunastylis spCaladenia prolata Calochilus robertsonii  Bunochilus sp non-fertile plant

Corysanthes diemenica

Did you get them?  Click on the image to go to the name and pictures of the flower. As a couple of the links are not working and until I have time to rectify them, click here to find the all answers in the Journal.
It will be on the last page.

More to come another time ………

2015 June Winning Photograph

06 sm PM Arachnorchis argocallaOf the five entries this month, four featured winter orchids. Lorraine Badger entered a Diplodium robustum, whilst Claire Chesson, Robert and Rosalie Lawrence all entered Urochilus sangineus. Though not the winning photographs it was interesting to see the differences between the U. sangineus with one being no taller than the small Acianthus pusillus next to it and another being taller than the rapier sedge.

But the winning photograph was the spring flowering Arachnorchis argocalla (White Beauty Spider Orchid) by Pauline Meyers. This is amongst our most threatened orchids and is dealt with in depth in the Recovery Plan For Twelve Threatened Orchids in the Lofty Block Region of South Australia 2010. This fungi dependent endemic orchid is rated Endangered both at State and National level.

Found in the Southern and Northern Lofty regions, it range has been severely reduced by possibly 80%. Since 1918 no plant has been found south of Adelaide.

Flowering from September to October, it is often found in grassy woodlands often growing on gentle southerly-facing hill slopes. The soil is a clay loam with a high humus content.

This beautiful orchid has one to two non-perfumed white flowers with thickened but not clubbed drooping lateral sepals and petals. The strongly recurved broad labellum is usually white, sometimes crimson, fringed with short teeth.

This is one of our larger spider orchids reaching a height of 60cms. The size of the plant flower and leaf help to distinguish it from other similar appearing orchids such as A. brumalis and albino flowers of A. behrii.

Like many of the spider orchids it takes 2 – 5 years to reach maturity and then has a potential reproductive life of 10 years. With an average pollination rate of less than 10%, the potential to increase the population is low and any threat to survival of the individual plants needs to taken seriously.

Some threats are obvious such as weed invasion including the garden escapees such as Topped lavender (Lavandula stoechas spp. stoechas) and action is being taken to curb the spread of weeds through targeted weeding programs.

Another threat is habitat loss. This has been the result of land clearing but sites are being protected either through conservation legislation or Heritage Agreements. Habitat loss can also occur indirectly and that is through Phytophthora being introduced into the sites. Although the direct effect of Phytophtora on the orchid is unknown, it is known that it can affect the plants that grow in association with this orchid. This threat can be reduced by all of us implementing good hygiene practices.

These were some of the threats noted in the Recovery Plan. This plan was not just defensive, ie attempt to halt and minimalize the damage; but it was also proactive with measures outlined to increase the population. These included seed and fungi collection eventually resulting in germination and cultivation with a view to re-introduction.

It is good to see that there is a plan and active steps are being taken to bring this orchid back from threat of extinction.

June 2015 other entrants
Photographers from L to R: Claire Chesson, Rosalie Lawrence, Lorraine Badger, Robert Lawrence

References

Websites accessed 1 July 2015

White Beauty Spider Orchid (Caladenia argocalla) Recovery Plan
http://www.environment.gov.au/archive/biodiversity/threatened/publications/recovery/c-argocalla/index.html
Caladenia argocalla – White-beauty spider-orchid, biodiversity species Profile and Threats Database
http://www.environment.gov.au/cgi-bin/sprat/public/publicspecies.pl?taxon_id=54991
Recovery Plan For twelve threatened Orchids in the Lofty Block Region of South Australia
http://www.environment.gov.au/system/files/resources/e362cfd2-a37b-443a-b007-db3a2b7b64dd/files/lofty-block-orchids-recovery-plan.pdf

Bates R J, South Australia’s Native Orchids 2011 DVD

2015 May Winning Photograph

05 sm PM Diuris hazeliae

Western Australia produces a lovely array of orchids and so it is not surprising to find in NOSSA photograph competitions that when a Western Australian species is entered it can often be the winner. This month was no different with Pauline Myers beautiful picture of a mass of Diuris hazeliae which was kindly identified by Andrew Brown.

This species has only recently been named in 2013 and as a result finding information was a challenge. Obviously there was no information in Jones Native Orchids of Australia (2006); and surprisingly the definitive Field guide to the Orchids of Western Australia (2013) A Brown et al did not appear to have any information.

But

  • the Western Australian Herbarium’s FloraBase (Western Australian Flora), has a map of distribution which is roughly a diagonal line from east of Geraldton to the north of Esperance. It is not listed as threatened.

    Distribution of Diuris hazeliae.  Map taken from the Western Australian Flora Base
    Distribution of Diuris hazeliae. Map taken from the Western Australian Flora Base
  • the Western Australian Herbarium lists the species as one of 59 new taxa added to their plant census in 2014.
  • from the Atlas of Living Australia it can be deduced that the flowering time is mainly August and September and is likely to be found in various types of shrublands margins including Eucalypt and mallee woodlands and appears to be mainly associated with rocky or granite outcrops.
  • and the National Species List APNI/APC yields the information that it was named after Hazel King, plant collector and conservationist with a special interest in orchids and was previously known by the phrase name Diuris ‘northern granite’ with a common name of Rosy-cheeked Donkey Orchid. It was found in a granite outcrop on her property Tampu (north of Beacon).

Fortunately Andrew Brown was able to help with extra information. It is listed in his book (page 212) but under the phrase name Diuris sp. Eastern Wheatbelt (Yellow Granite Donkey Orchid). Diuris sp. Northern Granite was found to be the same species and so the use of that name was discontinued but it does remain a synonym for Diuris hazeliae.

The following description is information updated from his book “Field Guide to the Orchids of Western Australia”

Diuris hazeliae D.L. Jones & C.J. French (yellow granite donkey orchid)

Flowering: August to September.

Description:

A common, inland donkey orchid 100 to 300 mm high with two to three basal leaves 50 to 150 mm long by 5 to 10 mm wide and up to seven predominantly yellow, brown marked flowers 20 to 40 mm across. Flowers are characterised by their broad petals, very broad dorsal sepal, narrow, reflexed, usually crossed lateral sepals and tri-lobed labellum with broad, spreading lateral lobes and a broad, flattened to convex mid lobe.

Distribution and habitat:

Found between Mullewa, Salmon Gums and Balladonia, growing in shallow soil pockets on granite outcrops and along drainage lines below rocky breakaways.

Notes:

Named in 2013 from specimens collected at Tampu, north of Beacon in September 1997. The species often forms very large colonies on granite outcrops.

Distinctive features:

Inland granite and breakaway habitat.

Very broad dorsal sepal.

Diuris hazeliae is part of the Diuris corymbosa complex of which, in 2013, there were only 10 of the 26 Western Australian species formally named. This situation has now changed with 14 now formally named. As a final word, Diuris orientis is South Australia’s only member of this complex.

More images of this species can be seen on Retired Aussies website http://www.retiredaussies.com/ColinsHome%20Page/OrchidsWA/Diuris/Diuris%20sp%20northen%20granite/Diuris%20sp%20northern%20granite.htm

 

References – All websites accessed on 29th May 2015-06-04

https://florabase.dpaw.wa.gov.au/science/nuytsia/755.pdf

https://florabase.dpaw.wa.gov.au/browse/profile/44161

https://biodiversity.org.au/nsl/services/api/instance/apni/772000

Jones, Native Orchids of Australia and its Territories (2006)

Brown, Field Guide to the Orchids of Western Australia (2013)